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LOLCats?
I'm thinking you have to go way back, something like lolcats, advice duck, ceiling cat, something where if you remove it from the equation meme culture just straight up does not develop, or does not develop the same way or at the same time.
Before LOLcats was Demotivational posters, which were just the generic motivational posters, but with a humorous twist. Motivational posters had a black border with text on the bottom, which made it easy to swap out in MS Paint with funny text back in the early Internet days. It was the first caption memes, which became even more popular when people started doing it with cute cat pics too.
I still have a large folder of black boxes. One day they'll be worth something!