Technology

Encrypted or not, the fact that someone else has it stored somewhere in their computers is dangerous.

Of course. You are simply over-representing this risk, though. Besides, regular people realistically don't need to worry about Proton being backdoored, because their device is 10-100x more likely to be breached instead. Security is not a binary, it's a shade. Performing a software update is also "dangerous". Do you check every time you update the software its code, to verify no malicious backdoor is there? No, exactly, you trust the maintainers and the package infrastructure.

The only recommended way to store private keys are offline and encrypted.

So you don't store them on your device(s) (encrypted)? I store my GPG keys that I use to sign software on my yubikeys. That said, email is something I check from my phone and multiple computers (as most people). Do you really use a hardware key to do on-the-fly decryption, every time someone sends you a message, from each device?

As a security engineer, I also generally discourage such absolute "recommendations". My threat model is different from a regular Joe threat model, and both are different from Snowden's. There is no such thing as "only recommended way", because this is not a religion, it's a risk decision. Most people use Gmail, where the content of their email is literally available server side. Those same people can gain privacy and security using GPG via Proton, and in their threat model "provider gets compromised and software backdoored" is completely irrelevant. Is it relevant in your threat model? Good, then yes, you should only store keys offline and encrypted. Actually, you shouldn't use email at all, and you should use dedicated tools and protocols that are meant for security, where metadata is not transmitted in clear text, for example. You should also have virtually no session duration and perform a full login with 2FA every time, you should probably access the software that you use to communicate only from a secure machine dedicated for the purpose etc..

I think you trust Proton a bit too much.

I simply have clear in my mind what my threat model is and what risks are acceptable. I perfectly fit in the "Anyone with privacy concerns" category in the threat model they built. What about you?